List of Publications


INNGJERDINGEN, K. T. THOLE, C. DIALLO, D. PAULSEN, B.S. HENSEL, A. Inhibition of Helicobacter pylori adhesion to human gastric adenocarcinoma epithelial cells by aqueous extracts and pectic polysaccharides from the roots of Cochlospermum tinctorium A. Rich. and Vernonia kotschyana  Sch. Bip. ex Walp, Submitted for publication

AUSTARHEIM, I., INNGJERDINGEN, K. T., PAULSEN, B. S., TOGOLA, A., DIAKITE, C. & DIALLO, D. 2013. Chromatographic immunoassays for Helicobacter pylori detection--are they reliable in Mali, West Africa? PAN African Medical Journal, 14:72. OPEN ACCESS

INNGJERDINGEN, K. T., LANGERUD, B. K., RASMUSSEN, H., OLSEN, T. K., AUSTARHEIM, I., GRONHAUG, T. E., AABERGE, I. S., DIALLO, D., PAULSEN, B. S. & MICHAELSEN, T. E. 2013. Pectic polysaccharides isolated from Malian medicinal plants protect against Streptococcus pneumoniae in a mouse pneumococcal infection model. Scand J Immunol, 77, 372-88.

TVETE INNGJERDINGEN, K., BALLO, N., ZHANG, B. Z., MALTERUD, K. E., MICHAELSEN, T. E., DIALLO, D. & SMESTAD PAULSEN, B. 2013. A comparison of bioactive aqueous extracts and polysaccharide fractions from roots of wild and cultivated Cochlospermum tinctorium A. Rich. Phytochemistry.


AUSTARHEIM, I., MAHAMANE, H., SANOGO, R., TOGOLA, A., KHALEDABADI, M., VESTRHEIM, A. C., INNGJERDINGEN, K. T., MICHAELSEN, T. E., DIALLO, D. & PAULSEN, B. S. 2012. Anti-ulcer polysaccharides from Cola cordifolia bark and leaves. Journal of Ethnopharmacology, 143, 221-7.

AUSTARHEIM, I., CHRISTENSEN, B. E., HEGNA, I. K., PETERSEN, B. O., DUUS, J. O., BYE, R., MICHAELSEN, T. E., DIALLO, D., INNGJERDINGEN, M. & PAULSEN, B. S. 2012. Chemical and biological characterization of pectin-like polysaccharides from the bark of the Malian medicinal tree Cola cordifolia. Carbohydrate Polymers, 89, 259-268.

AUSTARHEIM, I., NERGAARD, C. S., SANOGO, R., DIALLO, D. & PAULSEN, B. S. 2012. Inulin-rich fractions from Vernonia kotschyana roots have anti-ulcer activity. Journal of Ethnopharmacology, 144, 82-85.

INNGJERDINGEN, K. T., MESKINI, S., AUSTARHEIM, I., BALLO, N., INNGJERDINGEN, M., MICHAELSEN, T. E., DIALLO, D. & PAULSEN, B. S. 2012. Chemical and biological characterization of polysaccharides from wild and cultivated roots of Vernonia kotschyana. Journal of Ethnopharmacology, 139, 350-8.

LE, N. H., MALTERUD, K. E., DIALLO, D., PAULSEN, B. S., NERGARD, C. S. & WANGENSTEEN, H. 2012. Bioactive polyphenols in Ximenia americana and the traditional use among Malian healers. J Ethnopharmacol, 139, 858-62.

WILLCOX, M., SANOGO, R., DIAKITE, C., GIANI, S., PAULSEN, B. S. & DIALLO, D. 2012. Improved Traditional Medicines in Mali. Journal of Alternative and Complementary Medicine, 18, 212-220.


GLÆSERUD, S., GRONHAUG, T. E., MICHAELSEN, T. E., INNGJERDINGEN, M., BARSETT, H., DIALLO, D. & PAULSEN, B. S. 2011. Immunomodulating polysaccharides from leaves of the Malian medicinal tree Combretum glutinosum; structural differences between small and large leaves can substantiate the preference for small leaves by some healers. Journal of Medicinal Plants Research, 5, 2781–2790.

GRØNHAUG, T. E., KIYOHARA, H., SVEAASS, A., DIALLO, D., YAMADA, H. & PAULSEN, B. S. 2011. Beta-D-(1 -> 4)-galactan-containing side chains in RG-I regions of pectic polysaccharides from Biophytum petersianum Klotzsch contribute to expression of immunomodulating activity against intestinal Peyer's patch cells and macrophages. Phytochemistry, 72, 2139-2147.

PHAM, A. T., MALTERUD, K. E., PAULSEN, B. S., DIALLO, D. & WANGENSTEEN, H. 2011. DPPH radical scavenging and xanthine oxidase inhibitory activity of Terminalia macroptera leaves. Nat Prod Commun, 6, 1125-8.

PHAM, A. T., DVERGSNES, C., TOGOLA, A., WANGENSTEEN, H., DIALLO, D., PAULSEN, B. S. & MALTERUD, K. E. 2011. Terminalia macroptera, its current medicinal use and future perspectives. J Ethnopharmacol, 137, 1486-91.


GHILDYAL, P., GRØNHAUG TE, RUSTEN A, SKOGSRUD M, ROLSTAD B, DIALLO D, MICHAELSEN TE, INNGJERDINGEN M, PAULSEN BS. 2010. Chemical composition and immunological activities of polysaccharides isolated from the Malian medicinal plantSyzygium guineense. Journal of Pharmacology and Phytotherapy, 76-85.

GRONHAUG, T. E., GHILDYAL, P., BARSETT, H., MICHAELSEN, T. E., MORRIS, G., DIALLO, D., INNGJERDINGEN, M. & PAULSEN, B. S. 2010. Bioactive arabinogalactans from the leaves of Opilia celtidifolia Endl. ex Walp. (Opiliaceae). Glycobiology, 20, 1654-1664.

INNGJERDINGEN, K., NERGÅRD, C. S. INNGJERDINGEN, M., DIALLO, D., MICHAELSEN, T., Paulsen, B. S., Structure/activity relations of immunomodulating pectic polysaccharides from Malian medicinal plants. Abstracts of Papers, 239th ACS National Meeting, San Francisco, CA, United States, March 21-25.

SUTOVSKA, M., FRANOVA, S., SADLONOVA, V., GRONHAUG, T. E., DIALLO, D., PAULSEN, B. S. & CAPEK, P. 2010. The relationship between dose-dependent antitussive and bronchodilatory effects of Opilia celtidifolia polysaccharide and nitric oxide in guinea pigs. International Journal of Biological Macromolecules, 47, 508-513.



Togola, A., Hedding, B., Theis, A., Wangensteen, H., Rise, F., Paulsen, B.S., Diallo, D. & Malterud, K.E. (2009), 15-Lipoxygenase inhibitory effects of prenylated flavonoids from Erythrina senegalensis. Planta Med 75 (10), 1168-1170

Sutovská, M., Franová, S., Priseznaková, L., Nosálová, G., Togola, A., Diallo, D., Paulsen, B.S. & Capek, P. (2009), Antitussive activity of polysaccharides isolated from the Malian medicinal plants. Int. J. Biol. Macromol. 44 (3), 236-239



Gronhaug, T.E., Glaeserud, S., Skogsrud, M., Ballo, N., Bah, S., Diallo, D. & Paulsen, B.S. (2008), Ethnopharmacological survey of six medicinal plants from Mali, West-Afrca. J. Ethnobiol. Ethnomed. 4, 26

Inngjerdingen, M., Inngjerdingen, K.T., Patel, T., Allen, S., Chen, X., Rolstad, B., Morris, G.A., Harding, S.E., Michaelsen, T.E., Diallo, D. & Paulsen, B.S. (2008), Pectic polysaccharides from Biophytum petersianum Klotzsch, and their activation of macrophages and dendritic cells. Glycobiology 18(12), 1074-1084

Togola, A., Austarheim, I.M., Theis, A., Diallo, D. & Paulsen, B.S. (2008),Ethnopharmacological uses of Erythrina senegalensis; a comparison of three areas in Mali, and a link between traditional knowledge and modern biological science. J. Ethnobiol. Ethnomed. 4, 4:6

Togola, A., Inngjerdingen, M., Diallo, D., Barsett, H., Rolstad, B., Michaelsen, T.E. & Paulsen, B.S. (2008), Polysaccharides with complement fixing and macrophage stimulation activity from Opilia celtidifolia, isolation and partial characterisation. J. Ethnopharmacol. 115, 423-431

Togola, A., Naess, K., Diallo, D., Barsett, H., Michaelsen, T. & Paulsen, B. (2008), A polysaccharide with 40% mono-O-methylated monosaccharides from the bark of Cola cordifolia (Sterculiaceae), a medicinal tree from Mali (West Africa). Carbohydr. Polym. 73, 280-288



Bah, S., Jäger, A.K., Adsersen, A., Diallo, D. & Paulsen, B.S. (2007), Antiplasmodial and GABA(A)-benzodiazepine receptor binding activities of five plants used in traditional medicine in Mali, West Africa. J.Ethnopharmacol. 110(3), 451-457

Inngjerdingen, K.T., Kiyohara, H., Matsumoto, T., Petersen, D., Michaelsen, T.E., Diallo, D., Inngjerdingen, M., Yamada, H. & Paulsen, B.S. (2007), An immunomodulating pectic polymer from Glinus oppositifolius. Phytochemistry 68(7), 1046-1058

Inngjerdingen, K.T., Patel, T.R., Chen, X., Kenne, L., Allen, S., Morris, G.A., Harding, S.E., Matsumoto, T., Diallo, D., Yamada, H., Michaelsen, T.E., Inngjerdingen, M. & Paulsen, B.S. (2007), Immunological and structural properties of a pectic polymer from Glinus oppositifolius. Glycobiology 17, 1299-1310

Maiga, A.I., Malterud, K.E., Mathisen, G.H., Paulsen, R.E., Thomas-Oates, J., Bergstrø m, E., Reubsaet, J.L., Diallo, D. & Paulsen, B.S. (2007),Cell protective antioxidants from the root bark of Lannea velutina A. Rich., a Malian medicinal plant. J. Med. Plants Res. 1(4), 66-79



Bah, S., Diallo, D., Dembélé, S. & Paulsen, B.S. (2006), Ethnopharmacological survey of plants used for the treatment of schistosomiasis in Niono District, Mali. J. Ethnopharmacol. 105(3), 387-399

Bah, S., Paulsen, B.S., Diallo, D. & Johansen, H.T. (2006), Characterization of cysteine proteases in Malian medicinal plants. J. Ethnopharmacol. 107(2), 189-198

Inngjerdingen, K.T., Coulibaly, A., Diallo, D., Michaelsen, T.E. & Paulsen, B.S. (2006), A complement fixing polysaccharide from Biophytum petersianum Klotzsch, a medicinal plant from Mali, West Africa. Biomacromolecules 7(1), 48-53

Maiga, A., Malterud, K.E., Diallo, D. & Paulsen, B.S. (2006), Antioxidant and 15-lipoxygenase inhibitory activities of the Malian medicinal plants Diospyros abyssinica (Hiern) F.White (Enenaceae), Lannea velutina A Rich. (Anacardiaceae) and Crosspteryx febrifuga (Afzel) Bent. (Rubiaceae). J. Ethnopharmacol. 104(1-2), 66-79

Nergaard, C.S., Kiyohara, H., Reynolds, J.C., Thomas-Oates, J.E., Matsumoto, T., Yamada, H., Patel, T., Petersen, D., Michaelsen, T.E., Diallo, D. & Paulsen, B.S. (2006), Structures and structure-activity relationships of three mitogenic and complement fixing pectic arabinogalactans from the malian antiulcer plants Cochlospermum tinctorium A. Rich and Vernonia kotschyana Sch. Bip. ex Walp. Biomacromolecules 7(1), 71-79



Inngjerdingen, K.T., Debes, S.C., Inngjerdingen, M., Hokputsa, S., Harding, S.E., Rolstad, B., Michaelsen, T.E., Diallo, D. & Paulsen, B.S. (2005), Bioactive pectic polysaccharides from Glinus oppositifolius (L.) Aug. DC., a Malian medicinal plant, isolation and partial characterization. J. Ethnopharmacol. 101(1-3), 204-214

Maiga, A., Diallo, D., Bye, R. & Paulsen, B.S. (2005),Determination of some toxic and essential metal ions in medicinal and edible plants from Mali. J. Agric. Food. Chem. 53, 2316-1321

Maiga, A., Diallo, D., Fane, S., Sanogo, R., Paulsen, B.S. & Cisse, B. (2005), A survey of toxic plants on the market in the district of Bamako, Mali; Traditional knowledge compared with literature search of modern pharmacology and toxicology. J. Ethnopharmacol. 96(1-2), 183-193

Nergaard, C.S., Diallo, D., Inngjerdingen, K., Michaelsen, T.E., Matsumoto, T., Kiyohara, H., Yamada, H. & Paulsen, B.S. (2005), Medicinal use of Cochlospermum tinctorium in Mali Anti-ulcer-, radical scavenging- and immunomodulating activities of polymers in the aqueous extract of the roots. J. Ethnopharmacol. 96(1-2), 255-269

Nergaard, C.S., Kiyohara, H., Reynolds, J.C., Thomas-Oates, J.E., Matsumoto, T., Yamada, H., Michaelsen, T.E., Diallo, D. & Paulsen, B.S. (2005), Structure-immunomodulating activity relationships of a pectic arabinogalactan from Vernonia kotschyana Sch. Bip. ex Walp. Carbohydr. Res. 340(11), 1789-1801

Paulsen, B.S. & Barsett, H. (2005), Bioactive Pectic Polysaccharides. Adv. Polym. Sci. 186, 69-101

Togola, A., Diallo, D., Dembele, S., Barsett, H. & Paulsen, B.S. (2005), Ethnopharmacological survey of different uses of seven medicinal plants from Mali (West-Africa) in the regions Dioila, Kolokani and Siby. J. Ethnobiol. Ethnomed. 1(1), 7-15



Inngjerdingen, K., Nergaard, C.S., Diallo, D., Mounkoro, P.P. & Paulsen, B.S. (2004), An ethnopharmacological survey of plants used for wound healing in Dogonland, Mali, West Africa. J. Ethnopharmacol. 92(2-3), 233-244

Nergaard, C.S., Diallo, D., Michaelsen, T.E., Malterud, K.E., Kiyohara, H., Matsumoto, T., Yamada, H. & Paulsen, B.S. (2004), Isolation, partial characterisation and immunomodulating activities of polysaccharides from Vernonia kotschyana Sch. Bip. ex Walp. J. Ethnopharmacol. 91(1), 141-152



Diallo, D., Paulsen, B.S., Liljebäck, T.H.A. & Michaelsen, T.E. (2003), The malian medicinal plant Trichilia emetica; studies on polysaccharides with complement fixing ability. J. Ethnopharmacol. 84(2-3), 279-287



Diallo, D., Sogn, C., Samake, F.B., Paulsen, B.S., Michaelsen, T.E. & Keita, A. (2002), Wound Healing Plants in Mali, the Bamako Region. An ethnobotanical survey and Complement Fixation of Water Extracts from Selected Plants. Pharm. Biol. 40, 117-128

Mathisen, E., Diallo, D., Andersen, O.M. & Malterud, K.E. (2002), Antioxidants from the bark of Burkea africana, an African medicinal plant. Phytother. Res. 16, 148-153

Paulsen, B.S. (2002),Biologically active polysaccharides as possible lead compounds. Phytochem. Rev. 1, 379-387



Diallo, D., Marston, A., Terreaux, C., Toure, Y., Paulsen, B.S. & Hostettmann, K. (2001), Screening of malian medicinal plants for antifungal, larvicidal, molluscicidal, antioxidant and radical scavenging actvities. Phytother. Res. 15(5), 401-406

Diallo, D., Paulsen, B.S., Liljebäck, T.H. & Michaelsen, T.E. (2001), Polysaccharides from the roots of Entada africana Guill. et Perr., Mimosaceae, with complement fixing activity. J. Ethnopharmacol. 74(2), 159-171

Paulsen, B.S. (2001), Plant polysaccharides with immunostimulatory activities. Curr. Org. Chem. 5(9), 939-950



Diallo, D., Hveem, B., Mahmoud, M.A., Berge, G., Paulsen, B.S. & Maiga, A. (1999) , An Ethnobotanical survey of herbal drugs of Gourma district, Mali. Pharmaceutical Biology 37(1) , 80-91

Published Feb. 10, 2011 7:26 PM - Last modified Oct. 29, 2019 9:25 AM